Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark green, gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; disk-shaped, or star-like. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; dissected, or not dissected; palmately lobed; smooth margined. Epidermis waxy, or not waxy. Venation palmate. Secondary veins open; prominent. Smallest veins prominent; ladder like. Intramarginal vein absent. Stalk of blade more than half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at both ends. Stipules present, or absent (vestigial); free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia present; appearing as hair tufts; scattered along midrib. Basal glands present. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; terminal; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 2-3; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 15-20(-100) (many); filaments present; joined, or free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous, or triplostemonous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles solitary. Stigmas 2 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaves cuneate at base with two large glands; venation palmate; fruit large; the candlenut. Taxon images: aleurite.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.