Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark brown, black; not peeling in strips. Slash yellow, orange, red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; white; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves opposite (usually subopposite); not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units present; specialized inflorescence units umbels; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 1 (appearing as a ring); all joined; not fleshy; deciduous; not stalked; entire. Stamens 10-15; filaments present; free of each other; free of the perianth; all equal in size; triplostemonous, or polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3-5. Locules 3-5. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; about 1 cm long to much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaves opposite; flowers with long stalks, in a peduncled umbel; petals represented by a low ring. Taxon images: aetoxy.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.