Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; axillary; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across, or to about 10 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; 5; free, or all joined; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined, or free of each other; joined to the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3-5. Locules 3-10. Styles solitary. Stigmas 3-5 lobed (3 or 5); not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaves distichous; fruit with persistent large calyx and sometimes style. Taxon images: adinand.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.