Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash white, gray. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves opposite; not scale-like; simple, or compound; with three leaflets, or palmate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units present, or absent; specialized inflorescence units corymbs; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4; free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4; free; not fleshy; deciduous; not stalked; entire. Stamens 8; filaments present; free of each other; free of the perianth; of different sizes (alternating taller and shorter); diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 4. Locules 4. Styles solitary. Stigmas 4 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; splitting, or not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4-8 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaves uni- or trifoliate; flowers bisexual and 4-merous; filaments strongly tapering; fruit 4-locular. Taxon images: acro.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.